The upper molar cusps is relatively bunoselenomorph

He has a short and you can straight postprotocrista labially oriented. The fresh new preprotocrista, seemingly straight, meets a proper-set-up parastyle. This new metaconule is slightly faster and is huge into M3. The premetaconulecrista in addition to fossae are pretty straight forward as opposed to neocrista. The new rear fossa was angled, specifically with the M3. Out-of straight for the M1, the fresh new postmetaconulecrista will get sigmoidal on M3, missing this new metacone. The fresh labial cusps come in a comparable axis which have relatively upright cristae. The brand new paracone labial rib is outstanding, but as opposed to anterior groove. There’s absolutely no metacone labial rib. The fresh new para-, meso-, and metastyles is actually salient. The new lingual cingulum is quite weakened in order to missing like the entostyle.

The newest registered build of one’s next down premolar is normal for derived base pecoran and you can crown pecoran inside the that have a highly-put up mesolingual conid and posterior cristid , . More over, the current presence of a tooth retailer on p1 excludes an attribution to help you Prodremotherium otherwise Mosaicomeryx . Even if the pearly whites aren’t just like the bunoselenodont like in Amphitragulus elegans, new called specimens have quite primitive provides, such a highly faster metastylid, a minimal top, a tiny postentocristid, and although sideways compressed, bulged lingual cuspids to your all the way down molars. A few of these features prohibit an affiliation to help you often Dremotherium otherwise Bedenomeryx , . Furthermore, the latest really small-size away from “Amphitragulus” quercyi are a distinguishing ability for the most recent Oligocene ruminants out of Europe , , , .

Dremotherium cf. guthi away from Gaimersheim1: step one kept maxillary bones toward enamel row M1-M3 (SNSB-BSPG 1952II4855). off Mailhat: dos kept P4-M2 (MNHN LIM154). Babameryx engesseri out-of Gaimersheim1: step three left fragmentary mandible that have m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and (c) occlusal views. The size bar are dos cm.

Holotype introduced specimens pointers

Holotype. Jehenne (1987) has designated the specimen IPHEP LM.1969 because the holotype. This might be a limited right maxillary bone which have P3-M3 (, Fig. 2), from Los angeles Milloque (MP29, France).

Known specimens. Mailhat MNHN LIM154 left P4-M2 (this performs, Fig. six.2); Gaimersheim1 SNSB-BSPG 1952II4855 left maxillary which have M1-M3 (which performs, Fig. six.1); SNSB-BSPG 1952II4870 best m2.


Down long lasting dentition. The lower molar is quite defectively kept. However, which large ruminant possesses a fully selenomorph trigonid. The brand new cristid try elongated in addition to cuspids is actually large on the enamel slightly pleated. Brand new metastylid was well developed.

Top permanent dentition. The newest P4 try poorly maintained. The lingual region try shed. Brand new anterolabial cone possesses a deep failing labial rib. The fresh new prior stylid is broken. New rear you’re quick but expose. Zero neocrista blocks this new fossa. Brand new quadratic top molars during the occlusal check bear a well-put up metaconule quite smaller than this new protocone. The fresh new cusps have become selenomorph. This new preprotocrista therefore the postmetaconulecrista is actually labially based and a little upright. Brand new postprotocrista is a lot of time, very rounded, and labially built. It can be entirely separated (SNSB-BSPG 1952II4855) otherwise fused (MNHN LIM154) towards premetaconulecrista. The second comes to an end in the axis of your own mesostyle. Brand new rear basin can be keeps a little metaconule flex into the M2, big toward M3. This new paracone, possessing a good paracone rib, as well as the metacone, labially apartment and concave, try aimed. The brand new mesostyle and you may parastyle are well build. It means rounded small articles. The newest mesostyle is big out-of M1 so you can M3. The brand new entostyle try poor or absent. There can be neither a lingual cingulum encompassing the fresh protocone neither a great metastyle forming a properly-set up cuspid. But not, short anterior and posterior cingula exist.


That it ruminant is relatively complex in-being really-selenodont and achieving zero lingual cingula in upper molars, always found in Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and you may Babameryx , , , [61–63], . In comparison to Bachitherium, which kinds lacks an enthusiastic prior groove to the very big labial paracone rib , , . The fresh learned fossils become more just like the complex stem Pecora and the latest primitive crown Pecora (such as Amphitragulus, Dremotherium, Bedenomeryx). Brand new enamel is actually slightly pleated, whereas it is fundamentally effortless during the Prodremotherium. Also, the dimensions of such white teeth is a lot bigger than in Prodremotherium regarding the dated Quercy series and you will Pech Desse . The teeth become more selenomorph than in “Amphitragulus” and you may Bedenomeryx , , as well as the parastyle isn’t that anteriorly projected than just observed on the “Amphitragulus” feningrei , , . Taking a look at the neodiagnosis advised by the Mennecart , this type of fossils include all demonstrating options that come with Dremotherium. The fresh currently outlined varieties for the Europe are D. feignouxi (sorts of types) and you may D. guthi , , . The latter try slightly smaller than the type kinds , just like the fossils demonstrated right here. D. feignouxi simply happened for the Agenian (MN1-MN2; ), whereas D. guthi lived when you look at the latest Oligocene (MP28-MP30; ). But not, when you look at the lack of extra better-maintained matter, a decisive assignation towards the variety is not possible.